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What seems to have happened, is that the fighters who smiled were unintentionally leaking information about their own sense of weakness, so passing a psychological advantage to their opponent. The message from this is that taking control of your emotional state, avoiding smiling, and showing that you have the upper hand, even where that is not felt, is a good strategy in competitive environments.
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But, as the saying goes, if you can’t beat them, join them. Next time you're having your photograph taken, don't say ‘cheese’; say ‘cheeks’. Saying ‘cheeks will not only shape your mouth nicely, but will also remind you to squeeze your cheeks upwards into a visually satisfying, genuine-looking Duchenne smile.